Olli Astley got in touch with the Oxford Mail to say on Thursday at around 4.30pm he was heading southbound on the A34, near Kidlington, when he spotted the creature on the central reservation.
He described it as a 'puma/lynx-sized cat' which was 'black or dark brown'.
Mr Astley, who was with his 16-year-old daughter, added: "We haven't contacted the council but it had been removed overnight and by 8.15am this morning (Friday) as we went to take a picture.
He said he was 'sure' the animal was a cat due to its features but was far bigger than a normal pet feline.
The 47-year-old's description of the big cat is the latest in a long line of people going back to the 1990s to see what they swear is a big cat in west and north Oxfordshire.
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